Southern Division Portsmouth are in the semi-finals of the Women's Premier League Cup after a shock 2-1 extra time win over top-flight leaders Sunderland.
There will be a Northern Division club in the semi-final draw as Leicester City won 2-1 against West Ham United.
A new name will be on the Cup as Millwall Lionesses, the only previous winners still in the competition, lost 2-0 to Nottingham Forest.
Barnet, who won their quarter-final last month, are the last semi-finalist.
Last year's League Cup winners Leeds United were knocked out in the last round but returned to league action with a 3-0 victory over Reading.
Women's Premier League Cup
Portsmouth 2-1 Sunderland a.e.t.
Gemma Hillier's extra time winner took Pompey into the semi-finals for the first time in the club's history.
They had to come from behind to claim a famous victory, Kelly McDougall having given the Black Cats a 50th minute lead.
Justine Lorton equalised four minutes later and Hillier hit what proved to be the winning goal in the final minute of the first period of extra time.
Victory could have been more decisive but, in the closing stages, Lorton missed a penalty and Hillier struck the woodwork with a ferocious shot.
Millwall Lionesses 0-2 Nottingham Forest
Millwall's hopes of celebrating their 40th anniversary by lifting the League Cup for a second time, 14 years after their first triumph, were dashed by Forest.
Catherine Lawson gave the Premier League's second in the table team a 51st minute lead and Catherine Stainthorpe wrapped up victory seven minutes from time.
Leicester City 2-1 West Ham United
Leicester came from behind to win a physical battle of regional division sides with a late strike by Courtney Sweetman-Kirk.
Becky Merritt gave the visitors a 35th minute lead but Natalie Reay, who had earlier struck the post, equalised five minutes later.
Sweetman-Kirk then hit the Hammers' woodwork, but she found the net 12 minutes from time to take the Foxes through to the last four.
Women's Premier League
Leeds United 3-0 Reading
Leeds celebrated the return of manager Rick Passmoor with a much-needed victory that lifted them out of the relegation zone.
Passmoor, who resigned at the end of last season, has taken over the reins until the end of the current campaign following the sacking of Gemma Grainger.
The temporary boss saw his team take the lead five minutes after the interval through Tanya Rich's close range strike.
And goals in the last three minutes by Leah Galton and Carey Huegett brought a satisfying conclusion to Passmoor's first game back.
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