Wilkinson scored 27 points against Scotland on his comeback in February 2007 but the new-look England were still finding their feet during the Six Nations. England finished third after an encouraging win over France was sandwiched between dismal defeats by Ireland and Wales.
Ashton took England on tour to South Africa but club duties, rest, injuries and illness meant the party was missing 30 first-choicers.
And ahead of the World Cup pool game between the sides, South Africa laid down a massive marker.
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