Matt Crabb's 94th-minute winner earned victory as Eastbourne twice came from behind to win a thriller at New Lawn.
After a goalless first 32 minutes, six came in a mad burst before the break, sparked by Jamie Taylor's 30-yard lob.
Rovers replied with long-range strikes from Yan Klukowski and James Norwood, who was then sent off before three more goals in first-half injury time.
Taylor's 20-yarder levelled at 2-2, Ian Herring's penalty restored Rovers lead, before Richard Pacquette made it 3-3.
The penalty came following a foul on Ben Watson by Crabb, who made amends by crossing for top scorer Pacquette to head Eastbourne level.
After six goals in such a short space of time, the second half was never going to live up to the first.
But Eastbourne maintained the dramatic events of the night when they snatched victory deep into injury time, midfielder Crabb drilling home a loose ball for his first goal in seven games.
Forest Green:
Bittner, Bartlett, L Jones (Grimes 90), Caines, McDonald, Herring, M Fowler, Klukowski, Norwood, Watson, Styche. Subs Not Used: Gill, Davies, Else, Smith.
Sent Off: Norwood (42).
Booked: Bittner, Norwood, L Jones, M Fowler, McDonald.
Goals: Klukowski 35, Norwood 38, Herring 45 pen.
Eastbourne Borough:
Banks, Baker, Cook, Mambo, Brown, Brinkhurst (N Crabb 81), Weatherstone, Johnson, M Crabb, Taylor (Strevett 87), Pacquette. Subs Not Used: Smart, Austin, Atkin.
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